Ricinus communis

Castor bean, Castor Oil Plant

EuphorbiaceaeLeavesRootsSeeds/NutsFlowersPotential hazards — see below

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What to Eat

Edible parts: Leaves, Oil, CAUTION, POISONOUS, Root, Seeds, Flowers

The seed contains 35–55% of an edible oil used in cooking and is a rich source of phosphorus, 90% of which is in phytic form. Some caution is advised given the plant's toxicity.

Known Hazards

Ricinus is extremely allergenic, and has an OPALS allergy scale rating of 10 out of 10. The plant is also a very strong trigger for asthma, and allergies to Ricinus are commonplace and severe. The castor oil plant produces abundant amounts of very light pollen, which easily become airborne and can be inhaled into the lungs, triggering allergic reactions. The sap of the plant causes skin rashes. People who are allergic to the plant can also develop rashes from touching the leaves, flowers, or seeds. They can also have cross-allergic reactions to latex sap from the related Hevea brasiliensis plant. The toxicity of raw castor beans is due to the presence of ricin. Although the lethal dose in adults is considered to be four to eight seeds, reports of actual poisoning are relatively rare. According to the Guinness World Records, this is the world's most poisonous common plant. Ricin is also present in lower concentrations throughout the plant. If ricin is ingested, symptoms commonly begin within two to four hours, but may be delayed by up to 36 hours. These include a burning sensation in mouth and throat, abdominal pain, purging and bloody diarrhea. Within several days there is severe dehydration, a drop in blood pressure and a decrease in urine. Unless treated, death can be expected to occur within 3–5 days; however, in most cases a full recovery can be made. Poisoning occurs when animals, including humans, ingest broken castor beans or break the seed by chewing: intact seeds may pass through the digestive tract without releasing the toxin. The toxin provides the castor oil plant with some degree of natural protection from insect pests such as aphids. Ricin has been investigated for its potential use as an insecticide. Commercially available cold-pressed castor oil is not toxic to humans in normal doses, whether internal or external. As a non-freezing, antimicrobial, pressure-resistant lubricant for special purposes, either of latex or metals, or as a lubricating component of fuels. As sources of various chemical feedstocks. As a raw material for some varieties of biodiesel. As attractively patterned, low-cost personal adornments, such as non-durable necklaces and bracelets. Holes must not be drilled in the beans to make beads. The outer shell protects the wearer from the poison. Wearing castor beans has been known to cause rashes, and worse. As a component of many cosmetics. As an anti-microbial. The high percentage of ricinoleic acid residues in castor oil and its derivatives, inhibits many microbes, whether viral, bacterial or fungal. They accordingly are useful components of many ointments and similar preparations. As the major raw material (in oil form) for polyglycerol polyricinoleate, a modifier that improves the flow characteristics of cocoa butter in the manufacture of chocolate bars, and thereby reduces the costs. As a repellent for moles and voles in lawns.

Documented Toxic Effects

abdominal painacute kidney injurydehydrationdiarrheaepilepsygastrointestinal disorderhemolysishypotensionliver dysfunctionnauseaunconsciousnessvision disordersvomiting

Source: Encyclopedia of Life TraitBank. Toxicity often varies by plant part — consult the hazards section above for preparation-specific details.

Where to Find It

A tropical plant. It occurs throughout the country of Papua New Guinea up to about 2000 m altitude. It does well in humid areas but needs a well drained soil. It is tolerant to drought and light frost. It needs an open sunny position. It suits hardiness zones 9-12. In Yunnan.

Africa, Algeria, American Samoa, Angola, Argentina, Aruba, Asia, Australia, Balkans, Bangladesh, Belize, Benin, Bhutan, Botswana, Brazil, Britain, Bulgaria, Burkina Faso, Burundi, Cambodia, Cameroon, Canary Islands, Central Africa, Central African Republic, CAR, Central America, Central Asia, Chad, Chile, China, Colombia, Congo DR, Costa Rica, Côte d'Ivoire, Cuba, Cyprus, Djibouti, Dominican Republic, East Africa*, East Timor, Ecuador, Egypt, El Salvador, Eritrea, Eswatini, Ethiopia*, Europe, Fiji, France, Gabon, Ghana, Guiana, Guianas, Guinea, Guinée, Guinea-Bissau, Himalayas, India, Indochina, Indonesia, Iran, Iraq, Israel, Italy, Ivory Coast, Japan, Jordan, Kenya, Korea, Lebanon, Lesotho, Liberia, Libya, Luxembourg, Macedonia, Malawi, Malaysia, Maldives, Mali, Marquesas, Mauritania, Mediterranean, Mexico, Middle East, Morocco, Mozambique, Myanmar, Namibia, Nepal, New Caledonia, New Zealand, Niger, Nigeria, Niue, Norfolk Island, North Africa, North America, Northeastern India, Oman, Pacific, Pakistan, Palestine, Papua New Guinea, PNG, Portugal, Rwanda, Sahara, Samoa, Saudi Arabia, SE Asia, Senegal, Serbia, Seychelles, Sierra Leone, Somalia, South Africa, Southern Africa, South America, Spain, Sri Lanka, St Helena, Sudan, Suriname, Swaziland, Syria, Tajikistan, Tasmania, Tanzania, Thailand, Timor-Leste, Togo, Tunisia, Turkey, Türkiye, Uganda, United Arab Emirates, UAE, Uruguay, USA, Vanuatu, Venezuela, West Africa, West Indies, Yemen, Zambia, Zimbabwe,

Countries: Andorra, United Arab Emirates, Afghanistan, Antigua & Barbuda, Albania, Armenia, Angola, Argentina, American Samoa, Austria, Australia, Azerbaijan, Bosnia & Herzegovina, Barbados, Bangladesh, Belgium, Burkina Faso, Bulgaria, Bahrain, Burundi, Benin, Brunei, Bolivia, Brazil, Bahamas, Bhutan, Botswana, Belarus, Belize, Canada, Congo (DRC), Central African Republic, Congo (Republic), Switzerland, Cote d'Ivoire, Chile, Cameroon, China, Colombia, Costa Rica, Cuba, Cape Verde, Cyprus, Czechia, Germany, Djibouti, Denmark, Dominica, Dominican Republic, Algeria, Ecuador, Estonia, Egypt, Eritrea, Spain, Ethiopia, Finland, Fiji, Micronesia, France, Gabon, United Kingdom, Grenada, Georgia, French Guiana, Ghana, Gambia, Guinea, Equatorial Guinea, Greece, Guatemala, Guinea-Bissau, Guyana, Honduras, Croatia, Haiti, Hungary, Indonesia, Ireland, Israel, India, Iraq, Iran, Iceland, Italy, Jamaica, Jordan, Japan, Kenya, Kyrgyzstan, Cambodia, Kiribati, Comoros, St Kitts & Nevis, North Korea, South Korea, Kuwait, Kazakhstan, Laos, Lebanon, St Lucia, Liechtenstein, Sri Lanka, Liberia, Lesotho, Lithuania, Luxembourg, Latvia, Libya, Morocco, Monaco, Moldova, Montenegro, Madagascar, Marshall Islands, North Macedonia, Mali, Myanmar, Mongolia, Mauritania, Malta, Mauritius, Maldives, Malawi, Mexico, Malaysia, Mozambique, Namibia, New Caledonia, Niger, Nigeria, Nicaragua, Netherlands, Norway, Nepal, Nauru, Niue, New Zealand, Oman, Panama, Peru, French Polynesia, Papua New Guinea, Philippines, Pakistan, Poland, Puerto Rico, Portugal, Palau, Paraguay, Qatar, Romania, Serbia, Russia, Rwanda, Saudi Arabia, Solomon Islands, Seychelles, Sudan, Sweden, Singapore, Slovenia, Slovakia, Sierra Leone, San Marino, Senegal, Somalia, Suriname, South Sudan, Sao Tome & Principe, El Salvador, Syria, Eswatini, Chad, Togo, Thailand, Tajikistan, Timor-Leste, Turkmenistan, Tunisia, Tonga, Turkey, Trinidad & Tobago, Tuvalu, Taiwan, Tanzania, Ukraine, Uganda, United States, Uruguay, Uzbekistan, St Vincent, Venezuela, Vietnam, Vanuatu, Samoa, Yemen, South Africa, Zambia, Zimbabwe

How to Identify

A small evergreen tree. Often they are grown as annual plants. It grows up to about 6 m high and spreads to 2 m across. The stem is erect, stout and branching. The plant can form suckers. It has leaves with pointy lobes spreading out like fingers on a hand. The leaves are large and glossy. The leaves are on long leaf stalks. The flowers are red and large and woolly. Female flowers are at the top and male flowers lower down. Clusters of flowers produce seed capsules containing 3 spiny seeds. Several different cultivated varieties exist which are chosen for their leaf and flower colour.

How to Grow

Prefers a well-drained moisture retentive clay or sandy loam in full sun. Requires a rich soil and daytime temperatures above 20°c for the seedlings to grow well, though the seed may fail to set if temperatures rise above 38°C for an extended period. The plant requires 140 - 180 days of warm temperatures in the growing season in order to produce good crops of seed, and is readily killed by frost. The plant is reported to tolerate an annual precipitation in the range of 20 to 429cm, an annual temperature in the range of 7.0 to 27.8°C and a pH of 4.5 to 8.3. The castor-oil plant is a fast-growing shrub in the wild, reaching up to 12 metres in height, though it is much smaller when cultivated in the temperate zone. A very ornamental plant, although it is not winter hardy in Britain, it can be grown outdoors as an annual bedding plant for sub-tropical displays, and can flower and produce fruit in its first year in warm summers. It has been known to ripen a crop of seeds as far north as Christiana in Norway. Providing the plants water needs are met, yields of around 1 tonne per hectare have been achieved, with exceptional cases of up t 5 tonnes per hectare. It has a long history of cultivation as an oil-bearing and medicinal plant, having been grown in ancient Egypt. It is still widely cultivated for its seed in tropical and sub-tropical zones. There are many named varieties, some developed for ornamental use and others for oil production. Plants may need support in exposed areas.

Propagation: Sow seed in early spring in a warm greenhouse, in individual pots. When large enough to handle, prick seedlings out into individual pots and plant out after the last expected frosts. Seed retains viability for 2–3 years.

Medicinal Uses

Castor oil, extracted from the seed, is a well-established laxative with over 2,000 years of use. It is regarded as fast, safe, and gentle, typically producing a bowel movement within 3–5 hours, and is recommended for use in both the very young and the elderly. It is effective enough to be used routinely to clear the digestive tract in cases of poisoning. It should not be used for chronic constipation, where it addresses symptoms without treating the underlying cause. The flavour is somewhat unpleasant and may cause nausea in some individuals. The oil has a notable antidandruff effect and is well tolerated by the skin, making it a useful vehicle for medicinal and cosmetic preparations. When an alcoholic solution of castor oil is distilled in the presence of sodium salts of higher fatty acids, it congeals into a gel useful for treating non-inflammatory skin diseases and providing protection against occupational eczema and dermatitis. The seed is anthelmintic, cathartic, emollient, laxative, and purgative. It is rubbed on the temples for headache and powdered for application to abscesses and skin infections. In Tibetan medicine the seed is considered to have an acrid, bitter, and sweet taste with a heating potency, and is used for indigestion and as a purgative. A decoction of the leaves and roots is antitussive, discutient, and expectorant. Leaves are used as a poultice to relieve headaches and treat boils.

Other Uses

The seed contains 35–55% of a drying oil used not only in cooking but also as an ingredient in soaps, polishes, flypapers, paints, and varnishes, and as a lubricant, lighting fuel, and component of precision engine fuels. It is used to coat fabrics and protective coverings, in high-grade lubricant manufacture, in transparent typewriter and printing inks, and in textile dyeing — when converted into sulfonated castor oil (Turkey-Red Oil) it is used to dye cotton fabrics with alizarin. It is also used in producing 'Rilson', a polyamide nylon-type fibre. The dehydrated oil is an excellent drying agent comparable to tung oil, used in paints and varnishes. Hydrogenated castor oil is used in waxes, polishes, carbon paper, candles, and crayons. A fibre for rope-making is obtained from the stems; cellulose from the stems is also used for cardboard and paper. The growing plant is said to repel flies, mosquitoes, moles, and nibbling insects. The leaves have insecticidal properties.

Wikipedia

Source ↗

A fast-growing evergreen shrub reaching 1.5 m tall by 1 m wide. Hardy to UK zone 9, frost-tender. In leaf year-round, flowering July to September, seeds ripening September to November. Monoecious and self-fertile, wind-pollinated. Grows in light sandy, medium loamy, and heavy clay soils, prefers well-drained conditions, tolerates heavy clay. Suitable for mildly acid, neutral, and basic soils. Requires full sun and moist soil.

Production

Plants are fast growing. On poor soils plants produce flowers rather than leaves.

Other Information

A moderately common plant reportedly eaten in several places in Papua New Guinea. Normally it is considered very poisonous. It is cultivated.

Notes

There is only one Ricinus species. It can be invasive. The poison ricin in the seeds in soluble in water and not in the processed oil.

Names & Synonyms

Aamanaka, Ander, Arind, Boliyr, Bonobono, Butsu kasrael, Dan-khra, Dantha, Endaru, Guloo, Higuereta, Higuerilla, Jarak, Khrwia, Kobo, Lumono, Mahung, Mpuluka, Mufuta, Palma cristi, Reri, Ricino, Varenda, Veron, Woalime gol

Ricinus microcarpus PopovaRicinus megalosperma DelileRicinus perennis Steud.Ricinus purpurascens Bertol.Ricinus ruber Miq.Ricinus persicus PopovaRicinus minor Mill.Ricinus rugosus Mill.Ricinus medicus Forssk.Ricinus medius J. F. Gmel.Ricinus nanus Bald.Ricinus peltatus NoronhaRicinus messeniacus Heldr.Ricinus metallicus Pax & K. Hoffm.Ricinus spectabilis BlumeRicinus zanzibarinus PopovaRicinus rutilans Müll. Arg.Ricinus vulgaris GarsaultRicinus speciosus Burm. f.Ricinus urens Mill.Ricinus vulgaris Mill.Ricinus sanguineus Hort. ex GroenlandRicinus scaber Bertol.Ricinus tunisensis Desf.Ricinus undulatus BesserRicinus viridis Willd.Ricinus macrocarpus PopovaRicinus inermis Mill.Ricinus lividus Jacq.Ricinus laevis DC.Ricinus leucocarpus Bertol.Ricinus macrophyllus Bertol.Ricinus krappa Steud.Ricinus giganteus Pax & K. Hoffm.Ricinus glaucus Hoffmanns.Ricinus hybridus BesserRicinus japonicus Thunb.Croton spinosus L.Ricinus africanus Mill.Ricinus badius Rchb.Ricinus europaeus T. NeesRicinus armatus Andr.Ricinus digitatus NoronhaRicinus atropurpureus Pax & K. Hoffm.Ricinus borboniensis Pax & K. Hoffm.Ricinus cambodgensis BenaryRicinus angulatus Thunb.

References (80)

Achigan-Dako, E, et al (Eds), 2009, Catalogue of Traditional Vegetables in Benin. International Foundation for Science.
Addis, G., et al, 2005, Ethnobotanical Study of Edible Wild Plants in Some Selected Districts of Ethiopia. Human Ecology, Vol. 33, No. 1, pp. 83-118
Al-Sodany, Y. M., et al, 2013, Medicinal Plants in Saudi Arabia: I. Sarrwat Mountains at Taif, KSA. Academic Journal of Plant Sciences 6 (4): 134-145
Ambasta S.P. (Ed.), 2000, The Useful Plants of India. CSIR India. p 526
Aparicio, H., et al, 2021, Ethnobotanical study of medicinal and edible plants used in Nhamacoa area, Manica province-Mozambique. South African Journal of Botany 139 (2021) 318-328
Awas, T., 1997, A Study on the Ecology and Ethnobotany of Non-cultivated Food Plants and Wild Relatives of Cultivated crops in Gambella Region, Southwestern Ethiopia. Addis Ababa University. p 50
Blamey, M and Grey-Wilson, C., 2005, Wild flowers of the Mediterranean. A & C Black London. p 121
Bodkin, F., 1991, Encyclopedia Botanica. Cornstalk publishing, p 885
Bremness, L., 1994, Herbs. Collins Eyewitness Handbooks. Harper Collins. p 267
Brickell, C. (Ed.), 1999, The Royal Horticultural Society A-Z Encyclopedia of Garden Plants. Convent Garden Books. p 885
Burkill, H. M., 1985, The useful plants of west tropical Africa, Vol. 2. Kew.
Burkill, I.H., 1966, A Dictionary of the Economic Products of the Malay Peninsula. Ministry of Agriculture and Cooperatives, Kuala Lumpur, Malaysia. Vol 2 (I-Z) p 1944
Burnie, G.(Ed.), 2003, Annuals and Bulbs. The Gardener's Handbooks. Fog City Press. p 216
Cabalion, P. and Morat, P., 1983, Introduction le vegetation, la flore et aux noms vernaculaires de l'ile de Pentcoste (Vanuatu), In: Journal d'agriculture traditionnelle et de botanique appliquee JATBA Vol. 30, 3-4
Chowdery, T., et al, 2014, Wild edible plants of Uttar Dinajpur District, West Bengal. Life Science Leaflets. 47:pp 20-36 http://lifesciencesleaflets.ning.com
Chowdhury, M. & Mukherjee, R., 2012, Wild Edible Plants Consumed by Local Communities of Maldah of West Bengal, India. Indian J.Sci.Res.3(2) : 163-170
Cobley, L.S. (rev. Steele, W.M.) 2nd Ed., 1976, An Introduction to the Botany of Tropical Crops. Longmans. p 302
Cundall, P., (ed.), 2004, Gardening Australia: flora: the gardener's bible. ABC Books. p 1208
Davis, S.D., Heywood, V.H., & Hamilton, A.C. (eds), 1994, Centres of plant Diversity. WWF. Vol 2. p 340
Dharani, N., 2002, Field Guide to common Trees & Shrubs of East Africa. Struik. p 260
Demise, S. & Asfaw, Z., 2020, Ethno Botanical Study of Wild Edible Plants in Adola District, Southern, Ethiopia. International Journal of Research and Analytical Reviews (IJRAR) 7(2).
Ertug, F, Yenen Bitkiler. Resimli Türkiye Florası -I- Flora of Turkey - Ethnobotany supplement
Esperanca, M. J., 1988. Surviving in the wild. A glance at the wild plants and their uses. Vol. 1. p 196
Facciola, S., 1998, Cornucopia 2: a Source Book of Edible Plants. Kampong Publications, p 101
Flora of Australia Volume 49, Oceanic Islands 1, Australian Government Publishing Service, Canberra. (1994) p 232
Fowler, D. G., 2007, Zambian Plants: Their Vernacular Names and Uses. Kew. p 27
French, B.R., 1986, Food Plants of Papua New Guinea, A Compendium. Asia Pacific Science Foundation p 200
Ghosh, C. & Das A. P., 2011, Some useful and poisonous tea garden weeds from the Darjiling District of West Bengal, India. Pleione 5(1): 91 - 114
Hani Medicine of Xishuangbanna, 1999, p 407
Hearne, D.A., & Rance, S.J., 1975, Trees for Darwin and Northern Australia. AGPS, Canberra p 102
Hedrick, U.P., 1919, (Ed.), Sturtevant's edible plants of the world. p 571
Henty, E.E., 1980, Harmful Plants in Papua New Guinea. Botany Bulletin No 12. Division Botany, Lae, Papua New Guinea. p 61, Pl. 22
Hibbert, M., 2002, The Aussie Plant Finder 2002, Florilegium. p 252
Hide, R., et al, 1979, A checklist of some plants in the territory of the Sinasina Nimai (Simbai Province, Papua New Guinea), with notes on their uses. Department Anthropology, University of Aukland
Hossain, U. & Rahman, A., 2018, Study and quantitative analysis of wild vegetable floral diversity available in Barisal district, Bangladesh. Asian J. Med. Biol. Res. 2018, 4 (4), 362-371
http://palaeoworks.anu.edu.au/Nuno_PhD/04.pdf re Timor
http://www.botanic-gardens-ljubljana.com/en/plants
Hussey, B.M.J., Keighery, G.J., Cousens, R.D., Dodd, J., Lloyd, S.G., 1997, Western Weeds. A guide to the weeds of Western Australia. Plant Protection Society of Western Australia. p 146
Hu, Shiu-ying, 2005, Food Plants of China. The Chinese University Press. p 512
Jackes, B.R., 2001, Plants of the Tropics. Rainforest to Heath. An Identification Guide. James Cook University. p 54
Karim, F. M. & Dakheel, A, J., 2006, Salt-tolerant plants of the United Arab Emirates. 2006. International Center for Biosaline Agriculture, Dubai, UAE. p 86
Kew Plants of the World Online
Khan, M. & Hussain, S., 2014, Diversity of wild edible plants and flowering phenology of district Poonch (J & K) in the northwest Himalaya. Indian Journal of Sci, Res. 9(1): 032-038
Khanal, R., et al, 2014, Documenting abundance and use of underutilized plant species in the mid hill region of Nepal. ECOPRINT 21: 63-71, 2014
Kohila, A. & Kensa, V. M., 2019, Survey of Wild Edible Plants of Dhanakarkulam Panchayath, Tirunelveli District, Tamil Nadu, India. Kong. Res. J. 6(2): 20-27, 2019
Lamp, C & Collet F., 1989, Field Guide to Weeds in Australia. Inkata Press. p 239
Latham, P., 2004, Useful Plants of Bas-Congo province. Salvation Army & DFID p 239
Lazarides, M. & Hince, B., 1993, Handbook of Economic Plants of Australia, CSIRO. p 206
Lord, E.E., & Willis, J.H., 1999, Shrubs and Trees for Australian gardens. Lothian. p 259
Luizza, M. W., et al., 2013, Local Knowledge of Plants and their uses among Women in the Bale Mountains, Ethiopia. Ethnobotany Research & Applications 11:315-339
Marinelli, J. (Ed), 2004, Plant. DK. p 467
Martin, F.W. & Ruberte, R.M., 1979, Edible Leaves of the Tropics. Antillian College Press, Mayaguez, Puerto Rico. p 192
Mbuya, L.P., Msanga, H.P., Ruffo, C.K., Birnie, A & Tengnas, B., 1994, Useful Trees and Shrubs for Tanzania. Regional Soil Conservation Unit. Technical Handbook No 6. p 428
Milson. J., 2000, Trees and Shrubs of north-west Queensland. DPI p 76
Molla, A., Ethiopian Plant Names. http://www.ethiopic.com/aplants.htm
Obizoba, I. C. & Atu, L. N., 1993, Production and chemical evaluation of some food condiments of Nigeria. Plant Foods for Human Nutrition. 44:249-254
Ochse, J.J. et al, 1931, Vegetables of the Dutch East Indies. Asher reprint. p 288
Okigbo, B.N., Vegetables in Tropical Africa, in Opena, R.T. & Kyomo, M.L., 1990, Vegetable Research and development in SADCC countries. Asian Vegetable Research and development Centre. Taiwan. p 42
Paczkowska, G. & Chapman, A.R., 2000, The Western Australian Flora. A Descriptive Catalogue. Western Australian Herbarium. p 249
Peduruhewa, P. S., et al, 2021, Potential of Underutilized Wild Edible Plants as the Food for the Future – A Review. Journal of Food Security, Vol. 9, No. 4, 136-147
Peekel, P.G., 1984, (Translation E.E.Henty), Flora of the Bismarck Archipelago for Naturalists, Division of Botany, Lae, PNG. p 309
Pickering, H., & Roe, E., 2009, Wild Flowers of the Victoria Falls Area. Helen Pickering, London. p 63
Purseglove, J.W., 1968, Tropical Crops Dicotyledons, Longmans. p 180
Rashid, H. E., 1977, Geography of Bangladesh. Westview. p 290
Roa, J. A. G. & Boada, D. S. G., 2018, Fundación para el Fortalecimiento de la Fruticultura y Plantas Alimenticias no Convencionales en Colombia.
Sarma, H., et al, 2010, Updated Estimates of Wild Edible and Threatened Plants of Assam: A Meta-analysis. International Journal of Botany 6(4): 414-423
Selvam, V., 2007, Trees and shrubs of the Maldives. RAP Publication No. 2007/12 p 146
Singh, Dharampal, 1979, Castor, in Simmonds N.W.,(ed), Crop Plant Evolution. Longmans. London. p 84
Sujanapal, P., & Sankaran, K. V., 2016, Common Plants of Maldives. FAO & Kerala FRI, p 228
Terashima, H., et al, 1992, Ethnobotany of the Lega in the Tropical Rainforest of Eastern Zaire (Congo): Part Two, Zone de Walikale, African Study Monographs, Suppl. 19:1-60
Terra, G.J.A., 1973, Tropical Vegetables. Communication 54e Royal Tropical Institute, Amsterdam, p 70
Torre, de la, L., et al, 2008, Enciclopedia de las Plantas Útiles del Ecuador. Herbario QCA. Pontificia Universidad Católica del Ecuador. p 328
Tsering, J., et al, 2017, Ethnobotanical appraisal on wild edible plants used by the Monpa community of Arunchal Pradesh. Indian Journal of Traditional Knowledge. Vol 16(4), October 2017, pp 626-637
Uprety, Y., et al, 2012, Diversity of use and local knowledge of wild edible plant resources in Nepal. Journal of Ethnobotany and Ethnomedicine 8:16
van Wyk, Be, & Gericke, N., 2007, People's plants. A Guide to Useful Plants of Southern Africa. Briza. p 22
Vickery, M.L. and Vickery, B., 1979, Plant Products of Tropical Africa, Macmillan. p 32
Wiersema, J. H. & Leon, B., 2013, World Economic Plants. A Standard Reference CRC Press. 2nd Ed. p 592
Williams, C.N., Chew, W.Y., and Rajaratnam, J.A., 1989, Tree and Field Crops of the Wetter Regions of the Tropics. Longman, p 178
World Checklist of Useful Plant Species 2020. Royal Botanic Gardens, Kew
www.nationalherbarium.nl/thaueuph/ Flora of Thailand.